Doll was a four-year letterman and a 1948 All-Pacific Coast Conference halfback at USC, playing in the 1945 Rose Bowl, in which the Trojans beat Tennessee, and again in 1949, when they lost to Michigan. He led USC in rushing in 1944, '47 and '48 and ran for 1,067 career yards. He also led the team in receptions and scoring in '48.
Doll missed the 1945 season while serving in the Marines. USC said Doll was aboard the USS Missouri when the Japanese signed surrender papers on the ship to end World War II.
Doll was an assistant coach at the University of Washington in 1955, and he served in the same capacity at USC in 1957 and '58. Doll returned to the Lions as a coach in 1963 and '64 and again from 1978 to 1988. He was defensive coordinator for the Baltimore Colts in 1974 and for the Miami Dolphins in 1975 and '76.
Doll, a member of the Lions' 75th anniversary team selected two years ago, is survived by his wife, Diana; children Steven, Wendy, Kevan, Heidi and Michael; six grandchildren and one great grandchild.
The Associated Press contributed to this report.